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當(dāng)一名模仿殺手突然出現(xiàn)在拉斯維加斯時(shí)，拉塞爾和芬利在西雅圖工作時(shí)必須面對(duì)他們過去犯下的連環(huán)殺手案件。

　　第四季最后一集《A is for Answers》中，幾位少女一路追蹤，找到了愛麗森在費(fèi)城的藏身之所，她們向她坦白，認(rèn)為她的母親就是在莫娜卸下神秘人“A”身份后繼續(xù)折磨她們的繼任者。對(duì)于這一切愛麗森似乎并不特別意外，只是說局面太過復(fù)雜，并向她的伙伴吐露了內(nèi)心的秘密。

據(jù)聯(lián)邦調(diào)查局估算，目前美國(guó)境內(nèi)大約活躍著300名連環(huán)殺手。如果這些殺人惡魔有辦法相互溝通、聯(lián)系，甚至交流「謀殺經(jīng)驗(yàn)」，那會(huì)是怎樣一種情況？如果他們聯(lián)手犯案，甚至形成某種遍及全國(guó)的聯(lián)盟，那又會(huì)是怎樣一種結(jié)果？如果這些殺手中有一個(gè)變態(tài)殺人狂足夠聰明，聰明到能夠團(tuán)結(jié)所有的殺人惡魔，進(jìn)而激活一種人人效仿的「謀殺文化」，美國(guó)會(huì)變成什么樣子？這部由《吸血鬼日記》（The Vampire Diaries）制片人Kevin Williamson打造的驚悚罪案劇便以這三個(gè)疑問為故事背景。

　　偵探督察邁克·謝潑德擅長(zhǎng)自己的工作，卻不善于建立關(guān)系，到“布羅肯伍德”市破案時(shí)決定留下來，但很快謝潑德開始與克里斯汀·西姆斯偵探發(fā)生沖突。

　　As Leo's replacement Thomas Chamberlain arrives Nikki investigates the murders of Rachel Freedman, shot in the head, and her little autistic son Nathan, who was poisoned. Nikki rejects policewoman Anne Burchett's theory that Rachel killed them both but is perturbed when an impostor, claiming to be Rachel's sister, visits the morgue. At the same time the disfigured corpse of Eva Liron is discovered, killed with the same poison as Nathan. She had been the nanny to French hot shot footballer Isaac Dreyfus and was planning to blackmail him over a sex tape of them both. Jewish Isaac is a controversial figure due to his anti-Muslim sentiments but he denies ant part in the murders. Then another victim, Isaac's agent Mousa, is discovered dead.